The cambium is a lateral meristem that produces xylem cells to one side and phloem cells to the other to form the vascular system.

Photo of cellular plant material showing the cambium with xylem and phloem cells.

Divisions in the vascular cambium provide lateral growth to a stem.

The initial cell is called a fusiform initial and its periclinal division creates cells on both sides of the cambium line.

Photo of cellular plant material pointing out cambial cells.

In young stems, xylem and phloem appear in separated vascular bundles.

Cambium within the vascular bundle is called fascicular cambium, while the cambium between bundles is called interfasicular cambium.

Plant stem cross section showing vascular bundles and interfasicular cambium.

With age, the vascular system and the cambium forms a complete ring within the stem.

Older plant stem showing the xylem and cambium rings.

Secondary growth occurs in woody plants as the stems continue to expand.

Cross section of cellular plant material pointing out the ray, xylem, cambium, and phloem structures.

In addition to the vascular cambium, there is a cork cambium responsible for making bark or more correctly termed the periderm.

The periderm must continue to expand as the stem expands and the cork cambium is responsible for that axial growth.

Cross section of cellular plant material pointing out the periderm, cork cambium, and vascular cambium.

The cork cambium is also called the phellogen.

It gives rise to cells that form the periderm, which is the term for the protective tissue that replaces the epidermis in a stem.

Cross section of cellular plant material identifying the periderm.

The periderm is made of three layers.

  1. The outer phellem (or cork) layer.
  2. The middle phellogen (cork cambium).
  3. And the inner phelloderm, which is living parenchyma cells formed to the inner side of the phellogen.

Cross seciton of cellular plant material identifying the three layers of the periderm.